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Mecha Cheep are mechanical versions of Cheep Cheep that first appear in New Super Mario Bros. U. They have metal fins and a wind-up key at the back similar to that of Mechakoopas. In the game, Mecha Cheeps appear only in the level The Mighty Cannonship. They are impervious to Fireballs, but can be defeated with an Iceball or Star, or by leading a Targeting Ted into them.
Mecha Cheeps appear in Mario Kart 8 in the underwater section of Water Park and on posters seen in the entrance to the Sub Coaster at the beginning of the course, showing its structure. These posters say that the Mecha Cheeps have a built-in artificial intelligence system in their heads, and that their mouths function as inflow ports. The wind up key at the back provides the Mecha Cheep's propulsive power and says that they have levelling fins. Their steerage is apparently located in towards the tail in the underside of the fish. The poster also details the fuel efficiency as 888 mpg, and their serial number as 0026-2814.