From the Super Mario Wiki, the Mario encyclopedia
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Bristles are spiked monsters made of rock and are found in the game Paper Mario: The Thousand-Year Door. Similar to Clefts, Bristles have spiked tops, high defense, an immunity to fire attacks, and can be flipped over by an earthquake or an explosion. Unlike Clefts, Bristles do not lose their top-spiked defense when flipped and are also immune to close-range attacks like Gulp and Mario's Hammer, as their large spears pop out to protect against approachers. Bristles won't pop out their spears if asleep, frozen, immobilized, dizzy, poisoned, or confused, but they are still immune to direct contact moves such as Ms. Mowz's Kiss Thief like with Pokeys.
Despite their copious defenses, Bristles have very low HP and are easily taken out with POW Blocks, Earth Quakes, Superguards, or Earth Tremors. Mario is also capable of simply hammering them should he have the Spike Shield badge equipped, though without Power Smash, Piercing Blow, or the Ultra Hammer, it will do no damage.
Although Bristles have made no appearances outside of Paper Mario: The Thousand-Year Door, they are mentioned in Mario & Luigi: Partners in Time. Before the Mario Bros. learn the Bros. Ball move, Toadsworth the Younger tells his older self that he "looks like he has a Bristle in his britches".
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