From the Super Mario Wiki, the Mario encyclopedia
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Krashes are Kritters who ride in Mine Carts uncontrollably. They appear in Donkey Kong Country and Donkey Kong 64. In the latter game, Krashes were redesigned to have a muscular appearance and wield a club, both of which further distinguish them from Kritters and closer to Klubba from Donkey Kong Country 2: Diddy's Kong Quest.
Donkey Kong Country
In their debut appearance in Donkey Kong Country, Krashes appear in both Mine Cart levels, Mine Cart Carnage and Mine Cart Madness. In their first level appearance, Krashes ride a Mine Cart in the opposite direction of Donkey Kong and Diddy Kong, and if they ram into the Kongs, the lead one falls out of the Mine Cart. Although Krashes look vulnerable similarly to Kritters, the Kongs cannot jump their Mine Cart onto Krash, as this injures them while leaving the Krash unaffected. The Kongs must only avoid Krashes by jumping over them.
Krashes make their second and final appearance in Mine Cart Madness. Here, the Kongs jump out of their Mine Carts directly instead of carrying it with them. The Kongs can defeat Krashes very similarly to Kritters, by bouncing onto a Krash's head. Some Krashes sit idly at the start of a track railing, and the Kongs can jump on the Krash to hijack its Mine Cart and resume riding on the tracks.
In the Game Boy Advance remake, Krashes were changed to have a light purple skin.
Donkey Kong 64
In Donkey Kong 64, Krashes appear in the Mine Cart Rides where they ride on a track adjacent to the Kongs. Krashes were redesigned to have a muscular appearance and wield a club, both of which further distinguish them from Kritters. The Kongs can jump over the club to avoid taking damage. If the Kongs are hit, they lose three DK Coins.
Krashes are defeated if they crash into a pile of rocks or fall into a pit. The active Kong usually has to pull a lever to defeat Krash either way.