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Plessie first appears in World 1-4, Plessie's Plunging Falls, where Mario and company must ride it through a river to reach the Goal Pole. It also appears in World 5-1, Sunshine Seaside; World Bowser-2, Plessie's Dune Downhill, where it slides through sand; World Bowser-5, Deepwater Dungeon; and World Star-1, Rainbow Run. Plessie also makes a few cameos outside of these levels. In World Crown-Crown, Champion's Road, it appears at the end of the stage alongside other friendly non-playable characters gather to congratulate the player. It also appears in the game's title screen, and in the credits scene.
To control Plessie, players can tilt either the control stick or control pad, depending on the controller. To speed up and slow down, players tilt up and down respectively. Players can also press the jump button to jump, which can be used to defeat enemies and reach higher ground. If more players are riding Plessie, they can cooperate to tilt it further. If Plessie falls off a course, however, the lives lost is dependent on the amount of players riding it.
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