From the Super Mario Wiki
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Wheel Mice are enemies that appear in Wario World. They are Wario-sized mice that roll left and right in purple wheels in a few areas of Mirror Mansion. They appear out of narrow doorways in walls in lowered areas that Wario cannot enter, in infinite numbers. If the Wheel Mice's wheels roll over Wario, he is damaged. Wario can defeat them with punches, although this has little effect since more Wheel Mice will soon replace the defeated individuals. Empty wheels are also seen rolling by themselves in the same areas as Wheel Mice.