Princess Daisy (film character)
Daisy is an intelligent and analytical young woman who, perhaps in large part due to her saurian-heritage, is incredibly passionate about dinosaurs and paleontology. Even before discovering herself to be a dinosaur she felt a strong connection to dinosaurs and related to the idea of "trying to be human."
This passion later manifests itself as the growing desires of a "reptile brain" inspired by exposure to the parallel world. It is only her relationship with Luigi that grounds her humanity and prevents her from completely losing herself to those urges.
The film begins with Daisy's mother smuggling a metallic pod into the human dimension while being pursued by General Koopa. Koopa had previously slaughtered the Portal-Keeper ruling elite in a military coup, sparing only her due to his love for her. Knowing that he would force her to merge the dimensions she escaped and fled into the human world, making her way through the streets of New York City to the steps of a cathedral. She then deposits the pod on its steps, inserting a gleaming meteorite piece into it and frantically knocking on the door before fleeing again. An elderly nun opens the door just as she leaves and sees the pod, kneeling down to take it inside.
Daisy's mother then crosses the street, nearly being hit by a car before climbing down an open manhole that leads towards an abandoned subway shaft. She peers around in fear when Koopa suddenly steps from the shadows from behind and grabs her, demanding "the rock." Trying to escape, Daisy's mother causes a cave-in that both kills her and seals the tunnel, preventing Koopa from entering the human world to retrieve Daisy and the meteorite piece.
Back at the cathedral, several nuns surround the pod that now rests on an alter. The elderly nun from before removes the meteorite piece from it, causing the pod to open and reveal an egg inside. The egg begins to hatch and break apart until we see the infant Daisy, crying with life while her mother dies to save her. The elderly nun holds the meteorite piece up, which gleams as the scene cuts twenty years later.
Daisy is next seen as a paleontology student with a passion for dinosaurs that she doesn't quite understand. Her passion has brought her to work as the crew-leader at a project alongside the East River. However, the dig site is being hounded by Anthony Scapelli, a construction and plumbing mogul, because it had previously been the site of his company's construction project until the discovery of dinosaur bones forced him to step aside. Scapelli personally decides to visit the dig site in an attempt to settle his problem. Flanked by suited-goons, he demands to speak with whoever is in charge. This turns out to be Daisy who hops down from a ramp and asserts herself as "the boss." Unfazed by the girl, Scapelli sleazily eyes her body and threatens both her and the site if she doesn't end the digging soon. Worried, Daisy leaves another student in charge while she enters the city to call the university for more security.
Meanwhile, Spike and Iggy, Koopa's bumbling henchmen, sit in their car eating "dog" while discussing their failure to capture the right "girl." It is then that Iggy notices Daisy crossing the street and decide that she's the girl they have failed to locate. They begin to tail her, attempting to snatch her on the street only to be thwarted by their own stupidity. Daisy continues, unaware of the attempt on her life and arrives at a payphone currently being used by Luigi, who, stunned by her beauty, offers its use to her.
Unfortunately, Daisy soon finds that the university cannot increase their security. Dismayed, she approaches Luigi and his brother Mario to thank them for allowing her use of the phone. Mario, realizing his younger brother's infatuation with the girl, offers her a ride back to the dig site. Unsure of the offer, Daisy notices Spike and Iggy watching her from their car and accepts the brothers' offer, believing them safer than the two men she suspects are Scapelli's goons. The three talk along the way and get to know each other enough for Daisy to accept a dinner invitation from Luigi for later that night.
During that dinner Daisy reveals to Mario, Luigi, and Daniella that she was abandoned at birth and that she never removes the meteorite piece she wears around her neck as it is all she has of her past. Luigi admits that he also never knew his mother or father and was raised by his brother Mario, which prompts a shared gaze between the two and a moment of understanding. At Mario and Daniella's urging, Luigi offers to walk Daisy home following the meal.
During the walk home Luigi expresses embarrassment at his awkwardness. Daisy responds by explaining that she herself is embarrassed by her passion for dinosaurs, which she feels is an odd fascination to have. In the end, both are reassured by the others insecurity and smile. Daisy then offers to bring Luigi to the dig site to show him the fossils they have found.
The two race to the dig site, entering the tunnel that leads to the fossil pit. There's no security guarding the place, which Daisy explains is the reason she was using the phone earlier. She also explains to him how the fossils came to be found and the trouble Scapelli has been giving her crew. Luigi asks whether Daisy finds the underground chamber creepy. She says, to the contrary, it feels strangely "comfortable" to her.
The two begin to draw near for an intimate kiss, only to be interrupted by the sound of a nearby commotion followed by the sight of two of Scapelli's goons, Mike and Doug, running past them. The plumbing has been sabotaged in an attempt to flood the dig site and destroy the fossils. Daisy, aware that Luigi is a plumber, asks for his help.
After getting Mario, the two brothers begin to repair the plumbing while Daisy looks on. As they're about to finish, Iggy and Spike appear from behind, knocking them out and abducting Daisy. They pull her into the subterranean depths of the tunnel, leading her to the gateway between their world and the human world. Mario and Luigi, once the regain consciousness, follow. Finally, they come to what appears to be a dead end. However, they can hear Daisy's voice calling out to them from a seemingly solid rock wall. To their amazement, Daisy's face then appears through the rock wall as she struggles against Iggy and Spike. Luigi tries to grab her, but misses as she's pulled back and is left only with the meteorite piece worn around her neck.
Daisy is then pulled through the apathetic streets of Dinohattan before entering a cab. She is later seen being led into the holding cell where the four missing "Brooklyn Girls" are being held. It is there that she reunites with Daniella, who had been kidnapped only hours before. The presence of a familiar face comforts Daisy. She is then told by the Brooklyn Girls that they were kidnapped by someone known as "Koopa" who is seeking a princess. Daisy seems to suspect it is her, but says nothing. Later on in the movie, Lena and Koopa are having a mud bath. After Iggy and Spike come in, Koopa decides to see Daisy and orders Lena to do so. After going to the cell where Daisy is being held, Lena reveals that Daisy is in fact the Princess and reveals Daisy's past to her. Koopa also thinks that Daisy is the only one able to merge the dimensions and that is why the Brooklyn Girls have been kidnapped. While waiting for Koopa to meet her, Daisy meets Yoshi a dinosaur pet of Koopa. Despite her original fear of Yoshi, she slowly befriends the creature. Surprising her, Koopa comes in and after confirming everything Lena said, Daisy attempts to escape before being captured by a Goomba.
After Daisy is given food by a Goomba, Lena comes and tries to kill Daisy, but Yoshi pulls Lena back and Daisy escapes. While escaping, she runs into the Goomba from earlier and Spike and Iggy who are being dragged to their death. After igniting the Goomba, the trio escape after Daisy extinguishes the Goomba. Spike and Iggy claim they were two of her father's most loyal subjects and led here to Devo 4 where her father's fungal remains are. They then leave Daisy alone to process this. Yoshi comes in and Daisy pulls out the knife that Yoshi was stabbed by Lena with. After pulling the knife out, Yoshi motions Daisy towards a communications device. Using this, she communicates with the Mario Bros. and reveals her location. Once the Bros. come in, she shows them her father and directs Mario to where Daniella and the other Brooklyn Girls are. As Luigi and Daisy chase after Mario, they get caught by Koopa. When Mario and the Brooklyn Girls crash out of a vent, this distracts Koopa enough for Daisy and Luigi to escape. After attacking Koopa, the Bros. make him drop the meteorite. Lena grabs the meteorite piece and runs off. Luigi, Daisy, and the Brooklyn Girls proceed to chase Lena. Lena successfully inserts the meteorite despite the desperate pleas of Daisy. The resulting energy blast from the meteorite kills Lena. Daisy directs the Brooklyn Girls to the other dimensions so they warn those there of incoming invasion. Luigi and Daisy stay behind and successfully pull out the meteorite piece. The extinguished Goomba from earlier gives the two De-Evolution Guns. Big Bertha then gives Luigi a pair of Thwomp Stompers. The brothers then defeat Koopa and all of Dinohattan celebrates the end of their dictator. When Mario and Luigi decide to leave, they ask Daisy if she will be coming with them. Daisy then states she is going to stay here and spend time with her father. A few weeks later, she comes to the brothers' apartment and says she need their help with something. The three then go on another adventure.