From the Super Mario Wiki
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Lethiniums are flower-like enemies that appear in Kid Icarus: Uprising. They are very rare enemies and can fire a laser beam. They appear in Super Smash Bros. for Nintendo 3DS as Smash Run enemies. Like in the original game, they sit in the same spot, unable to flinch from attacks and can fire a laser at the player that can hit through the terrain. Shadowy enemies identical to Lethiniums also appear on the Master Fortress section of Classic in Super Smash Bros. for Wii U.
Super Smash Bros. for Nintendo 3DS