Parabones

Parabones are winged variants of Dry Bones and undead variants of Koopa Paratroopas. Like Dry Bones, they collapse when attacked and reanimate themselves moments later, though unlike Koopa Paratroopas, they usually do not lose their wings when stomped. They are the first and thus far only flying derived species of Koopa Troopa to appear in a 3D Super Mario platformer.

Super Mario 3D World / Super Mario 3D World + Bowser's Fury
Parabones first appear in Super Mario 3D World, replacing the normal Dry Bones in prior 3D games. They fly a small distance above the ground and charge into the player if they notice them. Like normal Dry Bones, if a Parabones is attacked without the use of a Super Star, Lucky Bell, or Invincibility Leaf, it will collapse temporarily, but reassemble itself after a short time; unlike their living counterparts, they do not lose their wings when they get hit. During its reassembly, it is vulnerable to getting attacked. A Parabones can also be permanently destroyed if it collapses over lava or a pit. They first appear in Fuzzy Time Mine. Parabones retain all these properties in the Nintendo Switch port, Super Mario 3D World + Bowser's Fury.

Super Mario Maker series
Parabones can be created in Super Mario Maker, Super Mario Maker for Nintendo 3DS and Super Mario Maker 2 by sticking wings to any Dry Bones, though like other enemies with wings attached to them, they are still referred to as the base enemy. While equipped with wings, they operate in the same manner as a red Koopa Paratroopa and, in the Super Mario World game style, they also throw bones at Mario. Unlike in Super Mario 3D World, stomping on a Parabones in this game removes its wings permanently, like Paratroopas. They no longer chase Mario and fly in a horizontal pattern to be consistent with the other styles.

Super Mario Odyssey
Parabones reappear in Super Mario Odyssey, where they appear in a secret area of Bowser's Kingdom and the Underground Moon Caverns in the Moon Kingdom. They wear white top hats and can be captured by using Cappy. As a Parabones, Mario can fly by pressing, and can shake the Joy-Con to fly faster. Like when Mario captures a Paragoomba, he cannot fly higher than where he started, but has no limits other than level boundaries when flying forwards. If Mario runs into lava or a Lava Geyser while flying, the Parabones disappears, though they respawn soon afterwards. The skull on the Caveman Headwear is stated to come from a Parabones.

Mario Tennis Aces
A Parabones acts as the referee of the Inferno Island court in Mario Tennis Aces.

Mario & Luigi: Bowser's Inside Story + Bowser Jr.'s Journey
Parabones appear in Mario & Luigi: Bowser's Inside Story + Bowser Jr.'s Journey as enemies and recruitable allies in the Bowser Jr.'s Journey mode, thus making their Mario & Luigi series debut. They are flying troopers, and are strong against Hammer Bros, Fire Bros, Boomerang Bros, and Ice Bros.

Skylanders: SuperChargers
In Skylanders: SuperChargers, Parabones appear as assistants to Hammer Slam Bowser, but only after the player took the Koopa Commander upgrade path and have purchased the Molten Bones Soul Gem upgrade. After he activates his Molten Monster form, he can attack one of his Koopa Paratroopas and turn them into Molten Parabones, which tend to do more damage and have more health than regular Paratroopas.

Paper Mario: The Origami King
Although Parabones do not appear physically in Paper Mario: The Origami King, concept art in Musée Champignon shows an origami Parabones. Additionally, their name is present in an unused string in the game's data.

Mario & Luigi: Bowser's Inside Story + Bowser Jr.'s Journey

 * Bowser Jr.'s Journey profile: Behold, the flying Dry Bones! For some reason, they're especially confident against red.

World of Nintendo figure
"Parabones are the reanimated bones of Paratroopas often found in the castles of Bowser or Haunted Mansions of the Mushroom Kingdom."